Islamic State insurgents stormed the historic Syrian city of Palmyra on Wednesday, fighting off pro-government forces who withdrew after evacuating most of the civilian population, state television said.

The capture of Palmyra is the first time the al-Qaida offshoot has taken control of a city directly from the Syrian army and allied forces, which have already lost ground in the northwest and south to other insurgent groups in recent weeks.

The central city, also known as Tadmur, is built alongside the remains of a oasis civilization whose colonnaded streets, temple and theater have stood for 2,000 years.