Twitter said on Thursday it had suspended hundreds of Russian-linked accounts and would ramp up enforcement of its spam rules as it probes online campaigns to influence the 2016 U.S. election.

Although the company's disclosures in briefings to U.S. congressional staff and a public blog post were its most detailed to date on the issue, Sen. Mark Warner, the top Democrat on the Senate Intelligence Committee, called the company's statements "deeply disappointing."

Warner, whose panel is investigating alleged Russian interference in the election, said Twitter officials had not answered many questions about the Russian use of the platform and that it was still subject to foreign manipulation.