The cause of India's deadliest train disaster in decades was linked to the signal system, the railway minister said Sunday, as families scoured hospitals and morgues for missing relatives with deaths expected to top 288.

Mounds of debris were piled high at the site of Friday night's crash near Balasore in the eastern state of Odisha, as workers started to clear the smashed carriages and the blood-stained wreckage where hundreds were also wounded.

Hospitals have been overwhelmed by the number of casualties.