Syrian army troops came closer to the Jordanian border on Thursday as the opposition and Russia agreed to resume suspended talks on a deal to end fighting that has sparked one of the fastest displacements in the course of the conflict.

Dark smoke rose over areas held by Syrian rebels near the border with Jordan as President Bashar Assad's Russian allies unleashed heavy airstrikes and government forces sought to advance on the ground.

The UNHCR refugee agency urged Jordan to open its borders to Syrians who have fled the fighting, saying the total number of displaced now stood at more than 320,000, with 60,000 of them gathered at the border crossing with Jordan.