U.S. President Donald Trump's former campaign manager, Paul Manafort, pleaded not guilty on Wednesday to a new indictment brought against him in the investigation into alleged Russian meddling in the 2016 election and will face trial in September.

Special Counsel Robert Mueller, who is running the Russia probe, is dialing up the legal pressure on Manafort, who has opted not to cooperate with investigators.

Manafort's former business partner, Rick Gates, another Trump ex-campaign official, decided last week to cooperate with the investigation.