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Marine biologist Osamu Shimomura, a co-winner of the 2008 Nobel Prize in chemistry, has died at the age of 90, Nagasaki University announced Sunday.

Shimomura, a graduate of the university and professor emeritus of Boston University, died of natural causes Friday in the city of Nagasaki. He won the Nobel Prize with Martin Chalfie and Roger Y. Tsien of the United States.

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